0000001 - Document - Cover Page link:presentationLink link:calculationLink link:definitionLink 9952151 - Statement - Condensed Consolidated Balance Sheets (Unaudited) link:presentationLink link:calculationLink link:definitionLink 9952152 - Statement - Condensed Consolidated Balance Sheets (Unaudited) (Parenthetical) link:presentationLink link:calculationLink link:definitionLink 9952153 - Statement - Condensed Consolidated Statements of Operations and Comprehensive Loss (Unaudited) link:presentationLink link:calculationLink link:definitionLink 9952154 - Statement - Condensed Consolidated Statements of Stockholders' Equity (Unaudited) link:presentationLink link:calculationLink link:definitionLink 9952155 - Statement - Condensed Consolidated Statements of Stockholders' Equity (Unaudited) (Parenthetical) link:presentationLink link:calculationLink link:definitionLink 9952156 - Statement - Condensed Consolidated Statements of Cash Flows (Unaudited) link:presentationLink link:calculationLink link:definitionLink 9952157 - Disclosure - Significant Accounting Policies link:presentationLink link:calculationLink link:definitionLink 9952158 - Disclosure - Fair Value Measurements link:presentationLink link:calculationLink link:definitionLink 9952159 - Disclosure - Marketable Securities link:presentationLink link:calculationLink link:definitionLink 9952160 - Disclosure - Intangible Assets link:presentationLink link:calculationLink link:definitionLink 9952161 - Disclosure - Research and Development Funding Collaboration Agreement link:presentationLink link:calculationLink link:definitionLink 9952162 - Disclosure - Collaboration Agreements link:presentationLink link:calculationLink link:definitionLink 9952163 - Disclosure - Balance Sheet Components link:presentationLink link:calculationLink link:definitionLink 9952164 - Disclosure - Liability Related to the Revenue Participation Right link:presentationLink link:calculationLink link:definitionLink 9952165 - Disclosure - Commitments and Contingencies link:presentationLink link:calculationLink link:definitionLink 9952166 - Disclosure - Common Stock link:presentationLink link:calculationLink link:definitionLink 9952167 - Disclosure - Stock-Based Awards link:presentationLink link:calculationLink link:definitionLink 9952168 - Disclosure - Net Loss Per Share link:presentationLink link:calculationLink link:definitionLink 9952169 - Disclosure - Non-Marketable Equity Investment link:presentationLink link:calculationLink link:definitionLink 9952170 - Disclosure - Segment Information link:presentationLink link:calculationLink link:definitionLink 9955511 - Disclosure - Significant Accounting Policies (Policies) link:presentationLink link:calculationLink link:definitionLink 9955512 - Disclosure - Fair Value Measurements (Tables) link:presentationLink link:calculationLink link:definitionLink 9955513 - Disclosure - Marketable Securities (Tables) link:presentationLink link:calculationLink link:definitionLink 9955514 - Disclosure - Collaboration Agreements (Tables) link:presentationLink link:calculationLink link:definitionLink 9955515 - Disclosure - Balance Sheet Components (Tables) link:presentationLink link:calculationLink link:definitionLink 9955516 - Disclosure - Commitments and Contingencies (Tables) link:presentationLink link:calculationLink link:definitionLink 9955517 - Disclosure - Stock-Based Awards (Tables) link:presentationLink link:calculationLink link:definitionLink 9955518 - Disclosure - Net Loss Per Share (Tables) link:presentationLink link:calculationLink link:definitionLink 9955519 - Disclosure - Segment Information (Tables) link:presentationLink link:calculationLink link:definitionLink 9955520 - Disclosure - Significant Accounting Policies (Details) link:presentationLink link:calculationLink link:definitionLink 9955521 - Disclosure - Fair Value Measurements - Schedule of Assets and Liabilities (Details) link:presentationLink link:calculationLink link:definitionLink 9955522 - Disclosure - Marketable Securities - Schedule of Available for Sale Securities (Details) link:presentationLink link:calculationLink link:definitionLink 9955523 - Disclosure - Marketable Securities - Narrative (Details) link:presentationLink link:calculationLink link:definitionLink 9955524 - Disclosure - Intangible Assets (Details) link:presentationLink link:calculationLink link:definitionLink 9955525 - Disclosure - Research and Development Funding Collaboration Agreement (Details) link:presentationLink link:calculationLink link:definitionLink 9955526 - Disclosure - Collaboration Agreements - Biogen (Details) link:presentationLink link:calculationLink link:definitionLink 9955527 - Disclosure - Collaboration Agreements - Sanofi (Details) link:presentationLink link:calculationLink link:definitionLink 9955528 - Disclosure - Collaboration Agreements - Takeda (Details) link:presentationLink link:calculationLink link:definitionLink 9955529 - Disclosure - Collaboration Agreements - Schedule of Cost Sharing Payments and Reimbursements (Details) link:presentationLink link:calculationLink link:definitionLink 9955530 - Disclosure - Balance Sheet Components - Schedule of Accrued Expenses and Other Current Liabilities (Details) link:presentationLink link:calculationLink link:definitionLink 9955531 - Disclosure - Liability Related to the Revenue Participation Right (Details) link:presentationLink link:calculationLink link:definitionLink 9955532 - Disclosure - Commitments and Contingencies - Schedule of Summarizes Lease Costs (Details) link:presentationLink link:calculationLink link:definitionLink 9955533 - Disclosure - Commitments and Contingencies - Narrative (Details) link:presentationLink link:calculationLink link:definitionLink 9955534 - Disclosure - Common Stock (Details) link:presentationLink link:calculationLink link:definitionLink 9955535 - Disclosure - Stock-Based Awards - Schedule of Stock Option Activity (Details) link:presentationLink link:calculationLink link:definitionLink 9955536 - Disclosure - Stock-Based Awards - Schedule of Assumptions Used for Estimating the Fair Value of Stock Options Granted (Details) link:presentationLink link:calculationLink link:definitionLink 9955537 - Disclosure - Stock-Based Awards - Schedule of Restricted Stock Activity (Details) link:presentationLink link:calculationLink link:definitionLink 9955538 - Disclosure - Stock-Based Awards -Schedule of Stock-Based Compensation Expense (Details) link:presentationLink link:calculationLink link:definitionLink 9955539 - Disclosure - Net Loss Per Share - Schedule of Computation of Basic and Diluted Net Loss Per Share (Details) link:presentationLink link:calculationLink link:definitionLink 9955540 - Disclosure - Net Loss Per Share - Narrative (Details) link:presentationLink link:calculationLink link:definitionLink 9955541 - Disclosure - Non-Marketable Equity Investment (Details) link:presentationLink link:calculationLink link:definitionLink 9955542 - Disclosure - Segment Information (Details) link:presentationLink link:calculationLink link:definitionLink